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Translated by W. Ogle.
144 pages - You are on Page 132
Of the two separate portions which constitute the head, namely the upper part and the lower jaw, the latter in man and in the viviparous quadrupeds moves not only upwards and downwards, but also from side to side; while in fishes, and birds and oviparous quadrupeds, the only movement is up and down. The reason is that this latter movement is the one required in biting and dividing food, while the lateral movement serve to reduce substances to a pulp. To such animals, therefore, as have grinder-teeth this lateral motion is of service; but to those animals that have no grinders it would be quite useless, and they are therefore invariably without it. For nature never makes anything that is superfluous. While in all other animals it is the lower jaw that is movable, in the river crocodile it is exceptionally the upper. This is because the feet in this creature are so excessively small as to be useless for seizing and holding prey; on which account nature has given it a mouth that can serve for these purposes in their stead. For that direction of motion which will give the greater force to a blow will be the more serviceable one in holding or in seizing prey; and a blow from above is always more forcible than one from below. Seeing, then, that both the prehension and the mastication of food are offices of the mouth, and that the former of these two is the more essential in an animal that has neither hands nor suitably formed feet, these crocodiles will derive greater benefit from a motion of the upper jaw downwards than from a motion of the lower jaw upwards. The same considerations explain why crabs also move the upper division of each claw and not the lower. For their claws are substitutes for hands, and so require to be suitable for the prehension of food, and not for its comminution; for such comminution and biting is the office of teeth. In crabs, then, and in such other animals as are able to seize their food in a leisurely manner, inasmuch as their mouth is not called on to perform its office while they are still in the water, the two functions are assigned to different parts, prehension to the hands or feet, biting and comminution of food to the mouth. But in crocodiles the mouth has been so framed by nature as to serve both purposes, the jaws being made to move in the manner just described.
Aristotle Complete Works
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