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Translated by W. Ogle.
144 pages - You are on Page 128
There are differences in the feet of quadrupeds. For in some of these animals there is a solid hoof, and in others a hoof cloven into two, and again in others a foot divided into many parts.
The hoof is solid when the body is large and the earthy matter present in great abundance; in which case the earth, instead of forming teeth and horns, is separated in the character of a nail, and being very abundant forms one continuous nail, that is a hoof, in place of several. This consumption of the earthy matter on the hoof explains why these animals, as a rule, have no huckle-bones; a second reason being that the presence of such a bone in the joint of the hind leg somewhat impedes its free motion. For extension and flexion can be made more rapidly in parts that have but one angle than in parts that have several. But the presence of a huckle-bone, as a connecting bolt, is the introduction as it were of a new limb-segment between the two ordinary ones. Such an addition adds to the weight of the foot, but renders the act of progression more secure. Thus it is that in such animals as have a hucklebone, it is only in the posterior and not in the anterior limbs that this bone is found. For the anterior limbs, moving as they do in advance of the others, require to be light and capable of ready flexion, whereas firmness and extensibility are what are wanted in the hind limbs. Moreover, a huckle-bone adds weight to the blow of a limb, and so renders it a suitable weapon of defence; and these animals all use their hind legs to protect themselves, kicking out with their heels against anything which annoys them. In the cloven-hoofed quadrupeds the lighter character of the hind legs admits of there being a huckle-bone; and the presence of the huckle-bone prevents them from having a solid hoof, the bony substance remaining in the joint, and therefore being deficient in the foot. As to the polydactylous quadrupeds, none of them have huckle-bones. For if they had they would not be polydactylous, but the divisions of the foot would only extend to that amount of its breadth which was covered by the huckle-bone. Thus it is that most of the animals that have huckle-bones are cloven-hoofed.
Aristotle Complete Works
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